Dorylus (Anomma) gribodoi Emery

Male Type locality Togo (Dorylus Gribodoi n. sp., Emery, 1892d: 570, illustrated, male) collected at Amu by Mr Gribodo see http://www.antweb.org/specimen.do?name=casent0903706
subspecies
confusus (Dorylus Gribodoi Em. var. confusa n. var., Santschi, 1915c: 246, male) from Ivory Coast, collected at Grand Bassam, by Lohier - no images on Antweb (July 2014)
insularis (Dorylus (Anomma) gribodi Em. v. insularis n. v., Santschi, 1937b: 98, male) from Fernando Po I. - see http://www.antweb.org/specimenImages.do?name=casent0911292
revisions and synonymy by Schöning et al. (2008)
junior synonyms (see lower page)
gerstaeckeri (Dorylus gerstäckeri n. sp., Emery, 1895j: 713, illustrated, holotype major worker only) from Ghana - type worker below
lamottei (Dorylus (Anomma) lamottei n. sp., Bernard, 1952: 219, illustrated, workers) from Guinea - type worker on linked page (see below)
all forms known .

My separation of the much redder form, with a more bulbous head on the major, as a distinct species, Dorylus (Anomma) quadratus (Santschi), has now (April, 2008) been confirmed by Schöning et al. (2008), although they synonymized quadratus with Dorylus (Anomma) kohli. Specimens that I recognise as kohli are much darker red-brown.

Emery's (1892d) description, with a drawing of the male head, is at . Emery (1895j) had a brief description, with an illustration; this is at . Santschi's (1937b) description of insularis is at .

Santschi (1915c) described the male of variety confusa as differing from the type in being darker, of uniform dark ferruginous brown; in gribodoi the thoracic sutures and the base of gastral segments has very distinct dark brown bands; in atratus the colour is black. The specimens were from Ivory Coast, Grand Bassam, by J.H. Lohier, 2 males, and from near Imbroko, by Posth, 1910, 3 males.

Schöning et al. (2008) synonymized D. gerstaeckeri and lamottei under Dorylus gribodoi, but they declined to separate it as in Anomma or Dorylus - http://www.mapress.com/zootaxa/2008/f/zt01749p052.pdf. Note that the single male and the workers were collected entirely separately but have been linked by genetic methods.

Wheeler (1922) gave records of gribodoi from Liberia (by Keiselbach), Ivory Coast (by A. Richard, Lohier, and at (D?)Imbroko by Posth), Ghana (no location), Nigeria (Lower Benue, by Lenfant), Congo and Zaïre.

From Guinea, Bernard (1952), calling it a purely western form, noted two typical males were taken from Mt. Nimba, one at Yalanzou, the other at Camp IV, 1000 m.

The photomontage is of a male from Taï, Ivory Coast, collected by Casper Schoning; collated from http://www.antweb.org/specimen.do?name=casent0172625

The holotype male can be seen at http://www.antweb.org/specimen.do?name=casent0903706

The photomontage is of a specimen from Taï, Ivory Coast, collected by Casper Schoning; collated from http://www.antweb.org/specimen.do?name=casent0172626&shot=p1&project=null

Dorylus (Anomma) gerstaeckeri Emery

Type locality Ghana (Dorylus gerstäckeri n. sp., Emery, 1895j: 713, illustrated, workers) from Accra .

Emery's (1895j) description is at . Emery's (1901c) description is at .

The original description was of a single worker, TL 8.0; but Emery (1901) received others from Cameroun which were longer, TL 10 mm. These were characterised by a square head about 1/3 of TL; vertex shiny and puncturation sparse; petiole slightly wider than long; overall body stocky (stockier than burmeisteri) and appendages shorter. Light brick-red, antennae ands legs yellowish, the head dunkel = darker or duller rust, the occiput darker

Raignier & van Boven (1955) under Dorylus (Anomma) emeryi note comparison with gerstaeckeri; emeryi being bigger TL 12.7 mm; head rectangular, slightly longer than wide, and slightly more than 1/3 of TL (in gerstaeckeri it is slightly less than 1/3).

The Ghana record was from Aburi (F. Silvestri, in Wheeler, 1922). One of the two species studied in Ghana during 1971 by Gotwald (1974). He found it only in the high forest areas, especially in and around cocoa at CRIG, where the almost exclusive prey was earthworms.

Also from Cameroun by Conradt (in Wheeler, 1922).

Specimens from Cameroun (Fotso Kuate) are shown in detail on Dorylus (Anomma) gerstaeckeri page.  Those match the Emery (1895j) description in form and colour, and are noticeably duller than the Schoning (Antweb) illustration; they are similar to the type gerstaeckeri but the form of the mandible suggests that morph for morph they are larger.

The photomontage is of a gerstaeckeri type worker collated from http://www.antweb.org/specimen.do?name=casent0903694

The holotype male can be seen at http://www.antweb.org/specimen.do?name=casent0903706

Oxford University Museum specimens

Dorylus (Anomma) gribodoi B Taylor det. (as lamottei)	Guinea T Humle Humle 01	20.ix.2001 Bossou 7°39'57 N 8°25'08 W		2
Dorylus (Anomma) gribodoi B Taylor det. (as lamottei)	Guinea T Humle Humle 05	8.ix.2001 Bossou 7°39'57 N 8°25'08 W		1
Dorylus (Anomma) gribodoi B Taylor det.	Benin J-F Vayssieres RVA 3052	20.iii.2013 Koro 09°22'21'' N 02°40'13'' E	Mangifera indica orchard prey of Oecophylla longinoda 1300-1400 h major	1
Dorylus (Anomma) gribodoi B Taylor det.	Cameroun A Fotso Kuate Dorylus sp1	17.v.2007 Matomb 03°54'30" N 11°25'58" E	Tuna bait in mixed crop fields	3

Dorylus (Anomma) lamottei Bernard

Type locality Guinea (Dorylus (Anomma) lamottei n. sp., Bernard, 1953b: 219, illustrated) from Mt. Nimba, 12 workers; 3 minors from Mount Tô; 4 medium workers, Nion crest, forest, 1300 m; 5 medium workers, Camp I, all collected by Lamotte; worker only known (see Bolton, 1995) .

Polymorphism - the photomontages are of specimens collected at Bossou, south eastern Guinea by Tatyana Humle (Humle 1, 20.ix.2001, and Humle 5, 8.ix.2001). The ant colonies were was observed as eaten by chimpanzees using the "ant-dipping" (using tools to gather the ants). The characters are quite distinctive, notably the short, near globose petiole with a spike-like subpetiolar process; the lighter colour and smaller size also seem consistent. See also the polymorphism montage at the bottom. It seems the species may be restricted to the Mt. Nimba massif and surrounding areas.

The various morphs are shown in detail on Dorylus (Anomma) lamottei morphs page.