Messor rufotestaceus (Foerster)

Iran list Type location Algeria (Formica rufotestacea, Foerster, 1850b: 489, worker; Messor rufotestaceus Foerst., Emery, 1908e: 439, illustrated full description, major & minor workers & queen; Forel, 1910a: male) - no type images on Antweb (June 2015)
junior synonyms
thoracica (Atta thoracica n. sp. Mayr, 1862: 742, illustrated, worker) from Syria - no images on Antweb (June 2015)
gracilinodis (Aphaenogaster gracilinodis n. sp., Emery, 1878b: 55, worker) from Syria - no images on Antweb (June 2015)

Perhaps not a Messor at all but as Emery (1875) had, an Aphaenogaster.

Foerster's (1850b) description is at . Emery (1908e) gave an illustrated (antenna) full description, this is at . Mayr's (1862) gave a description of thoracicus is at . Emery's (1878b) description of gracilinodis is at . Mohamed et al (2001) noted the underside of the head has only moderately curved and straight hairs.

Mayr's thoracicus illustration shows short sharp propodeal teeth; Emery (1908e) wrote of "stumpfen zahnen" = blunt teeth. Thome & Thome (1981b) showed merely angles at the dorsum and declivity junction.

Cagniant (1996a) in his key leads to two species which are essentially yellow-orange, have slender antennae and smooth gasters. One, Aphaenogaster foreli has propodeal spines, the other, which he gave as A. leveillei has no spines.  He noted the latter also as being synonymous with A. pallida laurenti of Santschi (1939d: 69). Significantly both were described from Algeria and appear to match Foerster's description. The Syria locations were added by Emery (1908) who also gave the dimorphism, something that Cagniant appears not to have seen in North African specimens.

Note: The difference and geographical distance between the Algeria type and the Syria-Iran forms suggests that two distinct species may be involved.  In the absence of type images it is impossible to resolve.

Egypt records - Wheeler & Mann (1916: 170) ; illustration in Tohmé & Tohmé (1981b); Wadi Gharandel, Sinai; - record in Mohamed et al (2001, illustrated). Sharaf list - material examined: El-Qosseima (North Sinai), 14.iii.1997 (4) M.R.Sharaf; Wadi Digla (Cairo), 13.iii.1955 (2); Wadi Digla (Cairo), 13.iii.1955 (1); Wadi Digla (Cairo), 13.iii.1955 (2); Wadi El-Arbaein, St. Catherine (South Sinai), 19.ii.1999 (1) M.R.Sharaf; Wadi Digla (Cairo), 13.iii.1955 (1) (ASUC).

Oxford University Museum specimens

Messor rufotestaceus B Taylor det.	Iran Torabi 36	06.v.2014 29.62245°N 52.582178°E 1578 m	Sadieh, Shiraz hand collected garden, rock; 14:15 h major & minor workers	2
Messor rufotestaceus B Taylor det.	Iran R Afshari 54	16.iv.2015 31.06°N 51.59°E 2066 m	Komeh, Semirom, Isfahan hand collected park, near river 1655h	1

The photomontage is of a major worker from Iran, Roshanak Torabi (36).

The photomontage is of a minor worker from Iran, RoshanakTorabi (36).

The photomontage of a worker from Iran is collated from http://www.antweb.org/specimen.do?name=cfh000017.