Cataulacus intrudens (F. Smith)

"intrudens" group: "rugosus" group Type location South Africa (Meranoplus intrudens, F. Smith, 1876d: 609, illustrated, all forms; placed in Cataulacus by Mayr, 1886c: 364) - queen see below
junior synonyms
batonga (Cataulacus Baumi For. r. Batonga n. stirps, Forel, 1913a: 114, worker) from Zimbabwe - see below
baumi (Catulacus Baumi n. sp., Forel, 1901d: 304, all forms) from Angola - see http://www.antweb.org/specimenImages.do?code=casent0904882
bulawayensis (Cataulacus Baumi For. r. Batonga For. v. bulawayensis, Forel, 1914d: 218; Arnold, 1917: 391, worker & queen) from Zimbabwe - see http://www.antweb.org/specimenImages.do?code=casent0909220
densipunctatus (Cataulacus johannae For. r. densipunctatus n. stirps, Stitz, 1923: 163, worker) from Namibia - no images on Antweb (September 2014)
foveosquamatus (Cataulacus foveosquamatus n. stirps, Santschi, 1937a: 58, illustrated, queen) from South Africa - see http://www.antweb.org/specimenImages.do?name=casent0912552
gazanus (Cataulacus baumi For. v. gazanus n. var., Santschi, 1928f: 208, worker) from Mozambique - see http://www.antweb.org/specimenImages.do?code=casent0912553
hararicus (Cataulacus hararicus n. spec., Forel, 1894b: 79, worker) from Ethiopia - see http://www.antweb.org/specimenImages.do?code=casent0909224
intermedius (Cataulacus intrudens Sm. stirps intermedius n. stirps, Santschi, 1917b: 287, worker) from Zimbabwe - see http://www.antweb.org/specimenImages.do?name=casent0912554
johannae (Cataulacus Johannae n. sp., Forel, 1895c: 250, worker & queen) from Madagascar - no images on Antweb (September 2014)
pseudotrema (Cataulacus baumi Forel stirps pseudotrema n. st., Santschi, 1926b: 244, worker) from Tanzania - see http://www.antweb.org/specimenImages.do?name=casent0912555
rugosus (Cataulacus intrudens Smith var. rugosus n. var., Forel, 1894b: 78, worker) from Mozambique - see http://www.antweb.org/specimenImages.do?name=casent0909234
subrugosus (Cataulacus rugosus For. var. subrugosus n. var., Santschi, 1914e: 26, worker; Santschi, 1937d: 236, queen) from South Africa (queen from Angola) - see http://www.antweb.org/specimenImages.do?name=casent0912557
umbilicatus (Cataulacus umbilicatus n. sp., Santschi, 1937a: 59, illustrated, queen) from Mozambique - see http://www.antweb.org/specimenImages.do?code=casent0900262
plus unavailable names
krugeri (Cataulacus rugosus For. krugeri var. nov. Prins, 1965d: 104, all forms) from South Africa - no images on Antweb (September 2014)
tangana (Cataulacus baumi Forel stirps pseudotrema Sants. v. tangana, Santschi, 1928f: 209, worker & queen) from Tanzania - see http://www.antweb.org/specimenImages.do?name=casent0912556
all forms known (in Bolton, 1974a: 43, illustrated) .

A group with fine or reduced sculpturation on the basal gaster - intrudens plus batonga, hararicus and intermedius -
Smith's (1876d) description of Meranoplus intrudens from South Africa is at - note that although Smith appeared to refer to an illustration of the worker, for comparison with the queen, his illustrations are of the queen and the male. A partial transcription (wrongly titled Smith, 1879) was given by Arnold (1917: 391), this is at . Forel's (1913a) description of batonga from Zimbabwe is at . An illustrated translation was given by Arnold (1917: 389), this is at . Forel's (1894b) description of hararicus from Ethiopia is at . Forel's (1895c) description of johannae from Madagascar is at . Santschi's (1917b) description of intermedius from Zimbabwe is at . A translation was given by Arnold (1920a: 403, 404), this is at .

Bolton (1974a: 42ff) synonymized the many names given above and drew attention to this species, which he describes as the most common of the genus in southern and eastern Africa, and appears to recognise two main groups (also separated as far as South Africa was concerned in a key by Arnold, 1920a: 403), this is at . Bolton's modern description (1974a) is at and .

The photomontage of the type queen is collated from http://www.antweb.org/specimen.do?name=casent0919586.
Bolton (1974a) related how he could not locate the F Smith queen in the BMNH and "this is presumed lost or destroyed". This specimen, however, clearly has all the right labelling, the original description being as Meranoplus intrudens. In his notes on ants in the BMNH collection, Mayr (1886c: 364) referred to Smith (1876) and that the Meranoplus intrudens belonged to Cataulacus. Mayr said that he had visited London in the summer of 1884. Curiously, Antweb does not seem to have found, or at least, photographed the BMNH specimen Bolton that designated as the lectotype worker.

The photomontage of the batonga cotype is collated from http://mcz-28168.oeb.harvard.edu/mcz/FMPro?-DB=Image.fm&-Lay=web&-Format=images.htm&Species_ID=29235&-Find

The photomontage of a specimen apparently wrongly identifed as wissmanni is collated from http://mcz-28168.oeb.harvard.edu/mcz/FMPro?-DB=Image.fm&-Lay=web&-Format=images.htm&Species_ID=29233&-Find

A group with coarse sculpturation on the basal gaster - rugosus plus baumi, bulawayensis, krugeri, pseudotrema, gazanus, subrugosus and tangana -
Forel's (1894b) description of rugosus from Mozambique is at . A translation was given by Arnold (1917: 392), this is at . Forel's (1901d) description of baumi from Angola is at . A translation was given by Arnold (1917: 388), this is at . Forel's (1914d: 218) description of bulawayensis from Zimbabwe is at . Arnold's (1917: 391, 1920a: 404) translation of bulawayensis is at . Santschi's (1914e) description of subrugosus from South Africa is at , Arnold's translation is on the rugosus card (above). Santschi's (1926b) description of pseudotrema from Tanzania is at . Stitz's (1923) description of densipunctatus from Namibia is at . Santschi's (1928f) description of gazanus from Mozambique is at . Santschi's (1928f) description of tangana from Tanzania is at . Prins's (1965b) description of krugeri from South Africa is at .

Smith (1876d) reported observations by J.M. Hutchison, at Weenen in South Africa, of the species making formicaria in the thorns of a species of acacia.

Queens only are known of - Santschi's (1937a) description of foveosquamosus from South Africa is at and his description of umbilicatus from Mozambique is at .

Oxford University Museum specimens

Cataulacus intrudens B Taylor det.	Tanzania G McGavin tree 5-55	13.i.1997 Mkomazi	ppkd collection from Acacia nilotica; 37M 0375378 UTM 9571003	2
Cataulacus intrudens B Taylor det.	Tanzania G McGavin tree 3-66	16.i.1996 Mkomazi 3°57.996' S 37°48.128' E	pkd collection from Lannea schweinfurthii	1
Cataulacus intrudens B Taylor det.	Tanzania G McGavin tree 3-67	16.i.1996 Mkomazi 3°57.996' S 37°48.128' E	pkd collection from Lannea schweinfurthii	4
Cataulacus intrudens B Taylor det.	Tanzania G McGavin tree 2-15	9.v.2007 Awae II 03°54'30" N 11°25'58" E	.	2
Cataulacus intrudens B Taylor det.	Namibia H Campbell Tree 09	07.ix.2009 Windhoek 23°13.836' S 18°24.097' E	Kuzikus Wildlife Reserve; 1340 m inside thorn on Acacia erioloba	1
Cataulacus intrudens B Taylor det.	Namibia H Campbell Tree 166	31.viii.2009 Windhoek 23°12.557' S 18°23'25" E	Kuzikus Wildlife Reserve; 1340 m inside thorn on Acacia erioloba	1

The photomontage is of a worker (hw 1.05 mm; dark scape; dark eyes) from Tanzania, Mkomazi Game Reserve, collector G McGavin (Tz 3-67). This appears close to the description of the variety tangana given by Santschi (1928f), type location Tanga, Tanzania.

The photomontage is of a large worker (HW 1.35 mm; orange scape; orange eyes) from Tanzania, Mkomazi Game Reserve; collector G McGavin (tz 5-55-3).

The photomontage is of a smaller worker (HW 1.1 mm) from Tanzania, Mkomazi Game Reserve; collector G McGavin (tz 5-55-3-2).

The photomontage of subrugosus is collated from The Smithsonian Institute cotype images at http://ripley.si.edu/ent/nmnhtypedb/public/specimeninfopage.cfm?publicconsumption=1&typespecimenID=759

Note: this is labelled as a cotype but was collected in Angola, Kambisa and Kampulu, Kasinga Region, vii.1932, 80 km S. of Vila-de Ponte (Santschi, 1937d); whereas the type location was in South Africa.

The photomontage is of a small worker (HW 0.8; orange scape; orange eyes) from Namibia, Mkomazi Game Reserve; collector H Campbell (Tree 09).

The photomontage is of a large worker (HW 1.35 mm; orange scape; dark eyes) from Namibia, Kuzikus Wild Life Reserve; collector H Campbell (Tree 166).