Pheidole minima Mayr
Type location Cameroun
(Pheidole minima nov. spec.,
Mayr, 1901a: 275, major & minor) collected at Mungo-Fluss,
collector R. Buchholz - no images available on Antweb (September 2014)
minima Mayr var. catella
n. var., Santschi, 1914d:
339, major & minor) from Nigeria (Olokemeji, collector F.
Silvestri - see below
minima Mayr stirps Faurei
n. st.., Santschi, 1920b:
15, minor & queen) from Gabon, Samkita , F Faure - see below
minima Mayr subspecies malelana
new subspecies, Wheeler,
1922: 133, major & minor, see below) from Zaïre (see
Bolton, 1995) - no images available on Antweb (September 2014)
major, minor and queen described .
Mayr's (1901a) description is at . Santschi's (1914d) description of catella
is at . Santschi's (1920b) description of faurei
is at .
TL major < 4.0 mm, minor ca. 2.0 mm (after Bernard,
Wheeler (1922) had catella also from Ghana
(Aburi, F. Silvestri). He gave notes on malelana, a very small
species, major TL 2.3 mm and worker TL 1.5 mm; major head as megacephala
in shape but mandibles no longer than broad, distinctly depressed on
the occipitum, see below. Both the major and workers generally seem to
be close to the form listed by me as Pheidole
nigeriensis (species B) except in generally lighter colour. It
was described from a single major and three workers nesting in a stem
of Hyphaene at Malela. The type form colouration was noted as
different, but he did not say in what way; Santschi had noted that catella
was of a colour similar to that now found in malelana but the
description was too brief for Wheeler to be sure of any comparison.
From Guinea, Bernard (1952) the finding of a
major and a worker at Nion crest, Station B6.26, 1300 m; which were
similar to the type form from Nigeria.
The Wheeler (1922) description of Pheidole minima
Mayr subspecies malelana was -
MAJOR - Length 2.3 mm. Head shaped much as in P.
megacephala, without the mandibles a little longer than broad,
distinctly but not broadly depressed in the occipital region. Eyes
small, flat, at the anterior third of the head. Clypeus flat,
ecarinate. Frontal area small, impressed; frontal carinae diverging,
reaching to the posterior third of the head, bounding distinct scrobes
for the antennal scapes, which are half as long as the head. Funicular
joints 2 to 8 distinctly broader than long, club longer than the
remainder of the funiculus. Mandibles large and convex, coarsely
bidentate at the tip. Thorax robust, pronotum very convex, with small
but distinct humeral tubercles. Mesonotum falling almost vertically to
the pronounced metanotal constriction, with a slight transverse
convexity in the middle. Propodeum broader than long, concave and
sloping in the middle, its spines rather erect, shorter than the
interval between their bases, with pointed tips. Petiole with rather
high, anteroposteriorly compressed, distinctly emarginate node.
Postpetiole only one and one-half times as broad as the petiole,
broader than long, with the sides angularly produced. Gaster much
smaller than the head, elliptical, convex, with subtruncate anterior
border. Legs stout, femora thickened in the middle.
Shining; mandibles sparsely punctate; clypeus rather smooth in the
middle, indistinctly rugulose on the sides; anterior two-thirds of head
with sharp, but not coarse, longitudinal rugae; occipital lobes with
small, sparse, piligerous punctures. Pronotum and gaster very smooth
and shining; pedicel smooth but less polished; mesonotum and propodeum
opaque, densely punctate. Hairs yellow, sparse, suberect on the body,
short and appressed on the legs and antennal scapes. Colour castaneous;
pronotum, first gastric segment, borders of clypeus, and mandibles
blackish; remainder of mandibles and clypeus, cheeks and anterior
portion of front, petiole and postpetiole yellowish red; legs brownish
yellow; terminal gastric segments pale brown; posterior borders of all
the gastric segments broadly yellowish.
WORKER - Length 1.5 mm. Head subrectangular, as
broad as long and as broad in front as behind, with very feebly convex
sides and nearly straight posterior border. Eyes just in front of the
middle. Mandibles with the entire apical border finely denticulate.
Clypeus convex, with rounded, entire anterior border. Antennal scapes
reaching beyond the posterior corners of the head to a distance equal
to twice their diameter. Thorax shaped much as in the soldier, but the
pronotum narrower and longer. Propodeal spines reduced to minute
slender teeth scarcely longer than broad at their bases. Superior
border of petiolar node straight and entire; postpetiole small, a
little broader than the petiole, subglobular. Pilosity, sculpture, and
colour as in the soldier, but the head smooth and shining, with only
the cheeks delicately longitudinally rugulose.
Described from a single soldier and three workers taken
by Lang from a colony nesting in a stem of Hyphaene at Malela,
Zaïre. This form agrees with the typical minima, in size and in
most of its characters but the colour is very different, the
postpetiole is much narrower in proportion to the petiole in both
soldier and worker; and the antennal scapes of the latter are decidedly
photomontage of the catella type major worker is
photomontage of the catella type
minor worker is
photomontage of the faurei type major worker is
photomontage of the faurei type minor worker is
Oxford University Museum
B Taylor det.
G Debout & A Dalecky
stipules of a Rubiaceae
B Taylor det.
Degraded secondary forest
518 m asl
B Taylor det.
|323 m asl; foraging
on palm leaves in later afternoon
photomontage is of specimens collected in Cameroun -
south-western tropical coastal forest area between Edéa and Campo
(McKey Wolbachia project) - Cameroon 31 from location KIEN
(Kienké - swamp/river alongside plantations at 2°52.02' N 9°59.13';
marshy land), 05 April 2001, in pouch-like stipules of a Rubiaceae.
These have the following characteristics -
Major - head in full-face view with only moderately
convex borders; in profile relatively slender with distinct acute
occipital junction. Mesostomal teeth well developed and anterior border
of ventral head capsule merged in darker strip with mesosomal border.
Scapes almost reaching occipital border; long but fine frontal carinae,
with longitudinal rugae laterally extending to occiput, main occipital
area smooth or almost so but distinct median line for full length of
face. Mandibles smooth, small basal tooth, edentulate border to strong
pair of apical teeth; clypeus ecarinate. Erect hairs on head long,
yellow-brown and moderately abundant. Pronotum domed, with fine
transverse rugae; fairly deep metanotal groove, propodeum dorsum
convex, declivity near flat, spiracle quite large, propodeal teeth
sharp and moderately long for group.
Minor - head
near-circular in full face view except
for almost straight anterior border (ignoring mandibles), unsculptured,
very shiny, with nuchal collar; eyes about 1/4 length of head, set just
forward of midline of face.
Minor worker from
Liberia, Yekepa; collector E Poiriet (Yekepa PF 10)
Minor worker from
Mali, Bamako; collector David King (King 43)