The Ants of Africa
Genus Camponotus subgenus Tanaemyrmex

Camponotus maculatus nominal varieties - forms separated as not being "maculatus" group species
Camponotus maculatus introduction

lacteipennis (Formica lacteipennis, Smith F., 1858b: 34, all forms) Camponotus lacteipennisSmith's (1858b) description is at {original description}. Major TL 9.5 mm, minor 8.5; base colour black; funiculi and legs rufo-testaceous, tarsi darker; head very large deeply impressed behind. Minor with elongate head. Port Natal. ? not a maculatus species (too small). Minor head similar to Camponotus caffer
Collected from http://www.antweb.org/specimenImages.do?code=casent0903512
South Africa
nubis (Camponotus (Myrmoturba) maculatus (Fabr.) nubis, subsp. nov., Weber, 1943c: 385, illustrated, Major & Minor)

{Camponotus maculatus nubis}Weber's (1943c) description is at {original description}.
Major TL 9.7 mm, no mention of any light patches. Baroni Urbani (1972: 129) appears not to have sighted the Weber type specimen but synonymized it with maculatus as a simple chromatic variety close to melanocnemis not with any obvious morphological differences. The minor specimen below obviously is fairly distinctive.  The small size suggests this is a distinct species.

No images on Antweb (October 2014).

South Sudan
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{short description of image}The photomontage of a wortker from South Sudan (as Bentoe, South Sudan Province), collected by Awatif Omer, 2006, Sudan 13 (right); exactly matches the alitrunk profile drawn by Weber, and has a very similar characters and colour.

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ugandensis
Camponotus liengmei ugandensis

sudanicus (Camponotus (Myrmoturba) maculatus (Fabr.), sudanicus subsp. nov., Weber, 1943c: 385, Major & Minor; synonymy by Baroni Urbani 1972: 125) {Camponotus maculatus sudanicus} Collated illustration (right) is of a cotype of Camponotus maculatus ssp sudanicus (Weber, 1943c: 385, major & minor workers; synonymy by Baroni Urbani 1972: 125) from South Sudan. The original photographs, together with enlarged images, are from the MCZ, Harvard University, website at - MCZ link. Weber's (1943c) description of sudanicus is at {original description}. Major TL 9.1 mm (AL 3.3 mm) body dark brown almost black; smaller than melanocnemis; darker than aegyptiacus and without spots; nest arboreal.
Minor tL 6.4 mm.
This does not seem to be any different to nubis see above. However, nubis was found nesting in the black montane humus under stones and among grass roots, whereas the single nest of sudanicus was in the stub of a tree branch. Despite describing both, Weber makes no reciprocal comparisons. The small size suggests this is a distinct species.
Sudan
Camponotus liengmei (Camponotus sexpunctatus n. spec. var. liengmei, Forel, 1894b: 67, Major & Minor; Forel, 1907g: 88, male) {Camponotus maculatus liengmei}Forel's (1894b) description is at {original description}. Arnold (1922: 636) gave a translation, this is at {original description}.
As with Camponotus sexpunctatus, this is a large form, major TL 15-17.3 mm; head elongate trapezoidal, distinctly longer than wide (liengmei shape); tibiae prismatic with setae present (piquants). The photomontage is of a cotype of Camponotus maculatus ssp liengmei. The original photographs, together with enlarged images, are from the MCZ, Harvard University, website at - MCZ link.
Given various statuses, this was placed as a subspecies/race of maculatus by Forel (1901h: 69), it was raised to species by Santschi (1923e: 292), then reduced to a junior synonym of maculatus by Baroni Urbani (1972: 125). The distinctive images of the cotype and a fresh specimen from Kenya, persuade me that it merits full species status.
Mozambique
minor worker {Camponotus maculatus liengmei} .
hansingi (Camponotus maculatus F. subsp. Liengmei Forel v. Hansingi, n. var., Forel, 1910e: 452, Major) Forel's (1910e) description is at {original description} Major only, TL 12-12.5 mm; head elongate trapezoidal, distinctly longer than wide (liengmei shape); tibiae prismatic with setae present (piquants). Beira, collector M H Hansing.  Probably associated or synonymous with Camponotus liengmei
No images on Antweb (October 2014)
Mozambique (also South Africa)
importunoides (Camponotus (Myrmoturba) maculatus F. r. Liengmei Forel v. importunoides, n. var., Forel, 1914d: 249, Major & Minor) Forel's (1914d) description is at {original description} TL 8-16.5 mm (a large major); minor head strongly narrowed behind (typical liengmei shape) (importunus not that shape) colour exactly as importunus. Probably associated or synonymous with Camponotus liengmei
No images on Antweb (October 2014)
South Africa
cognatomaculatus (Camponotus rubripes r cognatus var cognatomaculatus, Forel, 1886f: 151, worker; given as Forel, 1889: 255, unavailable name in Bolton 1995: 93) Forel's (1886f) mention of the name but without any obvious description (two specimens before him) is at {original description}. Essentially invalid. Kakoma - Equatorial Africa (there are several possibilities; the name Kakoma appears twice in Congo DR, possibly the most likely candidate
madecassa (Camponotus maculatus subsp. liocnemis var madecassa n.var., Emery, 1905d: 30, footnote, worker) Emery's (1905d) description is at {original description}. Major TL 13 mm, head 3.5 X 3, posterior tibia 3.5 mm. The form that Forel designated as C. maculatus collected by Grandidieri (Forel, 1891b: 1). See http://www.antweb.org/collection.do?name=ANTC3522. The specimens quite clearly are not C. maculatus. The major has a relatively longer, narrower head; the head has erect hairs on the underside of the head (also visible on the minor); the propodeum has a distinct angle between the dorsum and the steep declivity; there are no obvious setae on the ventral face of the tibiae. Madagascar
pessimus (Camponotus maculatus subspecies miserabilis Santschi variety pessimus, new variety, Wheeler, 1922: 235, Major & Minor)

Wheeler (1922) had - Major TL only 6-6.5 mm, Minor TL 5-5.5 mm, as miserabilis but much smaller; "the head of the major is distinctly narrower anteriorly, the cheeks being less convex and the frontal carinae are less approximated. Sculpture, pilosity and color very much as in miserabilis" - it seems possible that this is synonymous with Camponotus (Tanaemyrmex) agonius but that is inadequately described; image collated from http://mcz-28168.oeb.harvard.edu/mcz/FMPro?-DB=Image.fm&-Lay=web&-Format=images.htm&Species_ID=21488&-Find

Zare

{short description of image}
Contents Camponotus maculatus main entry
2010, 2013, 2014 - Brian Taylor CBiol FSB FRES
11, Grazingfield, Wilford, Nottingham, NG11 7FN, U.K.

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