|The Ants of
SUBFAMILY MYRMICINAE - Genus Pyramica
|Contents - Myrmicinae - MYRMICINAE Introduction|
In Tribe DACETINI
Dacetine Ants - Afrotropical Genera, after Bolton's revision of 1999, separable into those with elongate, linear mandibles, terminating in a fork of two or three spiniform teeth - Strumigenys, Microdaceton, Quadristruma - and those with basically triangular mandibles, some elongate but all without a terminal fork - Pyramica.
Diagnostic Features - Pedicel of two segments, postpetiole distinctly separated from the gaster, both segments with spongiform or lamelliform appendages of variable development. Antennae 6-segmented (4-segmented in a few instances), funiculus with two-segmented club. Antennal scrobes present. Mandibles basically triangular, even if elongated, and with a simple, single apical tooth. Roger's (1862a) genus definition is at - .
An early key to African and Madagascar dacetines is that of Santschi (1913b). It, with modern annotation, is at .
Bolton (1999) revived the Genus Pyramica Roger and synonymized all the following - Asketogenys, Chelystruma, Cladarogenys, Codiomyrmex, Codioxenus, Dorisidris, Dysedrognathus, Epitritus, Glamyromyrmex, Gymnomyrmex, Kyidris, Neostruma, Pentastruma, Serrastruma, Smithistruma, Tingimyrmex, and Trichoscapa - under this ancient Genus name. Those in bold have Afrotropical members.
Pyramica Roger (1862a: 251).
Type species: Pyramica gundlachi Roger, by monotypy, type location Cuba.
Pyramica junior synonym of Strumigenys, Roger (1863b: 40), Dalla Torre (1893: 145), Bingham (1903: 147).
Revived from synonymy as a subgenus of Strumigenys by Brown, W.L. (1948e: 110).
Junior synonym of Strumigenys (Brown, W.L. & Wilson, 1959b: 281), Brown, W.L. (1960c: 37).
Revived from synonymy by Bolton (1999).
Baroni Urbani & De Andrade (2007: 101) postulated that Strumigenys should be the senior synonym of (Afrotropical genera only given here): Cladarogenys, Codiomyrmex, Epitritus, Glamyromyrmex, Miccostruma, Pyramica, Serrastruma, Smithistruma and Trichoscapa. I note this but, as my key objective is the ease of identification, have retained the old genera with two main pages for Strumigenys and here Pyramica.
I find all these revisions by certain taxonomists confusing, to say
the least, and, although perhaps correct, simply irritating for those
with interests wider than narrow museum niceties. Thus, for now, I have
decided to regard the synonymised genera as subgenera and so preserve
recognition of species described in non-taxonomic research publications.
In compiling the key below and the subsidiary, linked, subgenera,
keys, I found that of Bolton (2000: 272 ff) limited in usefulness, at
least for anyone not equipped with a very high quality stereomicroscope
and able to mount tiny ants with, e.g. the mandibles open and the
ventral sides of the head visible. I have followed more simply visible
characters along the lines of the earlier Bolton (1983) keys.
Key to subgenera known from sub-Saharan Africa
|1||Mandibles elongate and linear||2|
|-||Mandibles relatively short||3|
|2||Antennal scapes with a broad, forward-produced, subbasal lobe, head with large orbicular hairs, clypeus with spatulate or otherwise broad hairs||subgenus Epitritus|
|-||Antennal scapes linear, head with only simple hairs, including clypeus||subgenus Cladarogenys|
|3||Mandible with a prominent basal lamella and no more than 17 denticles||4|
|-||Mandible without any lamella and with >20 denticles||subgenus Serrastruma|
|4||Fully closed mandibles with a distinct transverse border, bases overlapped by clypeus||5|
|-||When fully closed, mandibles with a strongly defined transverse border, with distinct gap between it and the clypeal margin; head without hairs; monotypic tramp||subgenus Trichoscapa|
|5||In profile mandibles increase in width from base to apex, with apex downcurved||subgenus Glamyromyrmex|
|-||In profile mandibles near parallel from base to apex, no more than extreme apex downcurved||subgenus Smithistruma|
© 2007, 2013, 2015 - Brian Taylor CBiol FSB FRES
11, Grazingfield, Wilford, Nottingham, NG11 7FN, U.K.