. |
Sexuals
only known |
. |
¤ |
Queen
and male only; queen with head black, shiny, rest of body dull
red-brown, appendages dark yellow; dorsum of body with clear
reticulation, but shiny with large puncturations; recumbent white
pilosity on head and body; TL 3.0-3.5 mm; ocelli small (B&F synonym
of punctatissima) |
Guinea
- breviceps |
¤ |
Queen only described; petiole scale with triangular
profile similar to transvaalensis but much smaller TL 3.2 mm;
ocelli relatively large (B&F synonym of dulcis) |
Sudan - muscicola |
¤ |
Male only; TL
ca 3.2 mm; uniform brown, ocelli large. The colour and gaster pilosity
resembles Hypoponera inaudax, which is
from the Congo Basin, but the large size suggests perhaps not (H. abellei has a male that is
smaller than the worker) |
Congo -
sp. undet. Hypoponera male |
. |
Workers |
. |
1 |
Dorsum
of propodeum separated from that of the mesonotum by a groove or
distinct suture - |
Hypoponera
- similoponera-group - 2 |
-- |
Dorsum
of mesonotum and propodeum forming a smooth unbroken surface without a
distinct suture - (true Hypoponera?) |
Hypoponera -
hypoponera-group - 28 |
. |
Hypoponera
- similoponera-group |
. |
2 |
Petiole scale thick, in
dorsal view triangular or at least less than square [similoponera-complex] |
3 |
-- |
Petiole scale thin, dorsally twice as wide
as long, in profile high and flat [squamoponera-complex] |
15 |
. |
similoponera-complex |
. |
3 |
TL
2.5 mm, yellow-brown, scape length surpassing the posterior border of
the head, eyes minimal and placed at about the centre of the cheeks |
Sudan -
ambigua |
-- |
Scape not reaching the occiput; eyes set
well forward |
4 |
4 |
Propodeum profile with
declivity equal in length to dorsum; generally brown, TL 2.5-3.0 mm;
eye of 3-5 facets |
5 |
-- |
Propodeum profile with declivity and
dorsum not equal in length, other features also different |
6 |
5 |
TL
2.5-3.0 mm; head with sides parallel, occiput slightly concave; eyes
3-5 facets; mandibles with 6 spaced out teeth, 3 long then 3 short; in
profile declivity of propodeum similar length to dorsum; head brown,
shiny; body dark red, gaster darker, tarsi yellow (B&F syn of punctatissima)
|
Pan-African
- cognata |
-- |
TL in
range 2.5-3.0 (2.7-3.5) mm; occiput convex; antennal scape not reaching
the occiput of the head; mandibles with 3 long anterior teeth, plus
12-15 shorter teeth; eye set well forward of head mid-length, with 3-5
facets; in profile propodeum with declivity and dorsum of similar
length; petiole scale thick, height less than three times the width;
subpetiolar process distinctively shaped with clear fovea; hind tarsus
with large pectinate spur and smaller spur; head black or rust-red, matt |
Pan-Mediterranean; tramp
- Ponera
coarctata |
. |
Propodeum
profile with declivity and dorsum not equal in length |
. |
6 |
TL
3.0 mm; propodeal declivity much longer than dorsum; antennal scape not
surpassing the occiput; eyes very close to the mandibles; mandibles
probably with less than 8 teeth; propodeum angle more obtuse than breviceps
(queen) colour brown |
Sudan -
mesoepinotalis |
-- |
Propodeum with declivity much shorter than
dorsum; elongated species, eyes with 0-2 facets |
7 |
7 |
Petiole scale remarkable
in its great volume, equal to that of the propodeum, very high and thick |
8 |
-- |
Petiole smaller not as large as propodeum;
mandibles with 5 teeth |
9 |
8 |
TL
2.6 mm; head elongated (HL 0.75, HW 0.50) and very convex, sides
straight but widening gently from the clypeus to a maximum just before
the occiput; mandibles slightly puncturated, with 4 large black teeth,
subtended by 5 smaller yellow teeth; clypeus slightly convex, with a
vague median carina, anterior margin rectilinear; eyes of a single
large, rounded facet, about one-sixth back from the mandibles; alitrunk
profile almost flat; propodeum short, rounded with declivity flat, a
little shorter than the dorsum, without marked lateral margins;
pilosity whitish-yellow, short and recumbent, densest on the tarsi;
head, propodeum and gaster densely puncturated with large punctures,
regularly spaced by twice their diameter; alitrunk with few punctures,
but with fine transverse rugae; entirely deep yellow, shiny, with only
the borders of the head brownish. |
Guinea
- petiolata |
-- |
TL
2.7-2.8; HL 1.2 X HW; mandibles with 8 denticles; propodeum apparently
very short; shiny, densely punctate; yellowish-brown, head darker |
Sudan - imatongica |
9 |
TL 3.5-3.8 mm; black,
appendages and gastral apex reddish |
South
Africa - sulcatinasis |
-- |
Shades of yellow to brown not black |
10 |
10 |
TL 3.7
mm; metanotal suture faint (not shown on drawing); head, alitrunk and
petiole dark yellowish-red, gaster rusty-red apex paler |
South
Africa - spei
|
-- |
TL < 3.5 mm |
11 |
11 |
Overall slender; petiole
in profile with an obtuse to flat dorsum; propodeum with near
right-angled transition from dorsum to declivity |
12 |
-- |
Stockier; petiole in profile with a
rounded summit; propodeum with angled but rounded transition from
dorsum to declivity |
14 |
12 |
Petiole in profile with
near vertical anterior and posterior faces so that summit no narrower
than base |
12A |
-- |
Petiole in profile with distinctly angled
faces so that summit narrower than the base |
13 |
12A |
TL
2.5-3.0 mm, head large, with convex sides; scape almost reaching
occiput; mandibles with 5 large, spaced out, teeth; eye of ca 4 facets;
penultimate segment of funiculus distinctly wider than long; first
segment of gaster shorter than second; dull red (yellowish) |
Seifert
characters - CS 616 CL/CW 1.172 SL/CS 0.773 PEW/CS 0.469 |
. |
 |
warm
temperate tramp species (possibly not in sub-Saharan Africa) - punctatissima |
-- |
Scape reaching no more than nine-tenths of
way back to occiput; penultimate segment of funiculus of equal width
and length; first segment of gaster longer than second segment (B&F
synonym of punctatissima)
|
Seifert characters - CS
558 CL/CW 1.181 SL/CS 0.734 PEW/CS 0.456 |
. |
 |
tropical tramp species - schauinslandi |
13 |
TL
2.4-2.8 mm; elongated species similar to but smaller than punctatissima
; mandibles with 4 large and one small teeth; propodeum declivity
shorter than dorsum; clypeus and alitrunk very shiny (dull in ssp);
yellowish-brown |
. |
. |
 |
tramp
species known from East Africa - ragusai |
-- |
TL
2.1 mm; HL 1.25 X HW; mandible with 9-10
teeth; eye position simply marked but no facets; propodeum declivity
flat and distinctly margined; shiny; yellow, head and gaster rust |
South Africa - traegaordhi |
. |
Stockier;
petiole in profile with a rounded summit; propodeum with angled but
rounded transition from dorsum to declivity |
. |
14 |
TL
2.7 mm; head rectangular; shorter mandibles & longer scapes almost
reaching occiput; eye of single facet; testaceous red with a brown spot
on front of head; appendages yellowish |
South
Africa - natalensis |
-- |
TL 2.4-2.8 mm; head
trapezoidal, narrower anteriorly; fairly deep metanotal suture; yellow
or slightly reddish; petiole node similar to that of Ponera
coarctata |
South
Africa - boerorum
|
. |
Hypoponera
- similoponera-group - squamoponera-complex,
petiole squamiform |
. |
15 |
Colour dark brown or
black |
16 |
-- |
Colour brown to yellow |
17 |
16 |
TL 3.0 mm; colour black;
appendages dark yellow (B&F synonym of eduardi)
|
South
Africa - dideroti
|
-- |
TL 5.5 mm; dark brown, head almost black |
. |
. |
 |
South Africa - transvaalensis
|
17 |
Scape short not reaching
occiput |
18 |
-- |
Scape longer even surpassing occiput |
23 |
18 |
Propodeum declivity
twice as long as dorsum; clypeus very short, without a carina, anterior
margin feebly arcuate |
19 |
-- |
Without the above combination of characters |
20 |
19 |
TL
4.0 mm; general features apparently similar to ursoïdea;
mandibles with 7 teeth; shiny, colour reddish
(B&F synonym of punctatissima)
|
Nigeria
- incisa |
-- |
TL
2.4-2.8 mm; head elongated, sides slightly convex, occiput concave;
eyes of 3-4 facets, placed about one-fifth of side back from mandibles;
mandibles with 3 anterior teeth, remainder denticles; antennal scape
not reaching the occiput by about its own width; generally matt, dull,
due to recumbent white hairs, dense over all the body. colour deep
red-brown, head even darker, appendages yellow-ochre
(B&F synonym of punctatissima)
|
Guinea - ursoïdea |
20 |
Colour clear brown,
anterior margin of clypeus feebly rounded; mandibles with 9-10
irregular denticles |
21 |
-- |
Colour yellowish or clear rust; mandibles
different |
22 |
21 |
TL 2.46
mm; head slightly elongated (HL 0.62, HW 0.5) with parallel sides; eyes
small, 2-3 facets; clypeus with short carina, not ridged in the middle;
dorsum of propodeum a little longer than the declivity, and with clear
angle at junction; shiny with relatively dense pilosity; dark
yellow-brown |
|
. |
 |
West
Africa & Gabon - lea |
-- |
TL 2.9 mm; eye of 5-6 facets; clypeus with feebly rounded
anterior margin, and a longer carina than lea, elongated by a
ridge; propodeum angle more rounded than lea; colour clear brown
(B&F synonym of punctatissima)
|
East Africa - petri |
22 |
TL 2.5; antennal scape
not reaching the posterior border of the head; declivity of propodeum
straight; reddish-yellow (B&F synonym of ragusae) |
eastern
Africa - aethiopica |
-- |
TL
2.2 mm; in profile propodeal declivity very sinuous; head relatively
short only slightly longer than wide, sides sub-parallel, widest in
anterior third; eyes distinctive, 3-4 facets, black, very elongated,
set some 1.5 times their length back from the mandibles; occiput
concave; antennal scapes not reaching the occiput; shiny, with little
puncturation; fine whitish hairs; testaceous yellow, gaster darker
except anteriorly |
Guinea - sinuosa |
. |
Scapes
reaching or surpassing occiput |
. |
23 |
Propodeal declivity
twice as long as the dorsum, with transition smoothly rounded; although
declivitous face distnctly bordered |
24 |
-- |
Propodeal declivity only slightly longer
than the dorsum |
25 |
24 |
TL
2.7-3.0 mm; scapes surpassing the occiput; clypeus rectilinear, or
feebly concave apically; mandibles with 3-4 clear teeth, and 9-10
reduced denticles; colour yellow-brown to yellow-red |
. |
. |
 |
eastern
Africa, Gabon & ?Guinea - dulcis |
-- |
TL 2.6
mm; antennal scape surpassing the occiput; eye of one facet, close to
the mandibles, which have only feeble teeth; clypeus margin with an
obtuse central tooth; propodeum profile similar to that of dulcis;
colour brown |
. |
. |
 |
Sudan & Congo - lotti |
25 |
Petiole scale with
narrow rearwardly curved profile |
26 |
-- |
Petiole scale with wholly vertical profile |
27 |
26 |
TL
2.5-2.7 mm; head elongated (HL 0.75, HW 0.48), sides somewhat convex,
widest towards the occiput, which is concave; eye with 3-4 facets,
sited about one-fifth of length from anterior; scape slightly
surpassing the occiput; clypeus strongly convex, with a limited carina,
anterior margin straight or feebly concave; mandibles with 5 clear
teeth, the last two small, preceded by 7-9 more or less distinct
denticles; yellow pubescence all over, moderately dense except on the
gaster; head and petiole dark brown, alitrunk and gaster redder
appendages brownish-yellow (B&F synonym of dulcis) |
Guinea
- lamottei |
-- |
TL
2.5-2.7; head moderately elongated (HL 0.76, HW 0.55) with subparallel
sides, occiput near straight; scape clearly passing the occiput; eye
black and very distinct, even though of a single large facet; placed
about one-sixth of way back from mandibles; clypeus short, nearly flat,
little puncturated, anterior margin straight; mandibles distinct, being
slender and puncturated, with 12-13 teeth of which 4 are large and
inequal; less puncturation than sinuosa and petiolata;
on head and gaster dense but very fine puncturation; generally shiny,
with little pilosity. colour entirely testaceous
(B&F synonym of dulcis)
|
Guinea - mandibularis |
27 |
Petiole
scale in profile tapering and narrow even at base; no description given |
. |
. |
 |
Cameroun,
Gabon & Fernando Po - rothkirchi
|
-- |
TL
1.7 mm; head 1.25 longer than wide, sides straight; anterior margin of
clypeus slightly arcuate; petiole scale in profile quite thick
narrowing only at apex; yellow, appendages paler |
Guinea - angustata |
-- |
Hypoponera - hypoponera-group |
-- |
28 |
Petiole
squamiform; eyes rounded, 6 facets; placed about one quarter back from
mandibles;
TL 2.2 mm; head shortish (HL 0.75, HW 0.58), sides feebly convex,
largest in anterior third; occiput concave, slightly surpassed by the
antennal scapes; mandibles a little more elongated than other
relatives; distally with 3 large teeth, then 3 shorter teeth, subtended
by 13 denticles; clypeus very distinct, very short, with a prominent
central triangular plate, extended centrally with a median tooth;
alitrunk profile very flat; declivity of propodeum twice as long as
dorsum, flat with an arched face, limited by fine margins, not dilated
as wings; abundant white pilosity on the gaster, less dense elsewhere;
dorsum very finely but distinctly puncturate; matt testaceous-yellow;
paler on the head; pronotum, petiole, femora and antennae darker dirty
yellow (B&F synonym of dulcis) |
Guinea
- villiersi |
-- |
Eyes reduced to no more than 2 facets;
petiole scale a thick scale or node |
29 |
29 |
TL 1.8-2.2 mm; eyeless;
propodeal declivity with no more than slight bordering or rounded;
generally yellow; petiole scale in profile narrowing dorsally; (once
all regarded as races of coeca) |
30 |
-- |
TL 2.6-3.3 mm; eyes with 1-2 facets or
position visible; petiole scale thick; colour dark yellow to brown |
32 |
. |
TL
1.8-2.2 mm, generally yellow and eyeless |
. |
30 |
TL 1.8-2.0 mm; dorsal profile of thorax rectilinear, with
no lateral waisting between mesonotum and propodeum; dark yellow-brown
(B&F synonym of coeca) |
Cameroun
- myrmicariae |
-- |
From above sides of alitrunk with distinct
narrowing of posterior mesonotum to propodeum |
31 |
31 |
TL ca.
2.2 mm; occiput narrower than maximum width, scapes not reaching the
occiput, mandibles with 6 teeth; anterior 5 larger, clypeus without an
apical point; petiole scale triangular; pale yellow, testaceous |
.
|
.
|
Specimen
from Gabon (as
with inaudax?
below)
|
West
Africa & Congo Basin - coeca |
-- |
TL in range 1.8-2.2
mm; mandible with only 5 visible teeth
- otherwise as for coeca of which it may be a variety; colour
dark reddish-yellow |
Congo Basin - inaudax |
. |
TL at
least 2.6 mm |
. |
32 |
TL 2.0-2.8 mm; generally
slender with low body profile; petiole scale thick with a domed dorsum
profile; propodeum with a vertical, slightly concave declivity that is
shorter than the dorsum |
33 |
-- |
TL 2.6-3.3 mm; eyes nil or around 2
facets; petiole scale thick and vertical |
34 |
33 |
TL
2.3-2.5 mm; HL 1.5 X HW; mandibles with 4 large teeth and 7-12
denticles; yellow-red, appendages yellower |
. |
. |
 |
North
African - abeillei |
-- |
TL
2.0-2.8 mm; eye distinct - similar to abeillei
but head slightly shorter and funiculus with distinct club (may be
synonymous with abeillei) |
Ethiopia - orba |
-- |
TL
2.0-2.5 mm; head wider (HL 0.47, HW 0.39); petiole scale with dome apex
in profile; shiny with relatively dense pilosity; yellow-brown,
appendages paler |
. |
. |
 |
West
Africa - cammerunensis |
- |
TL 2.6-3.3 mm; eyes
nil or around 2 facets; petiole scale thick and vertical |
- |
34 |
Eyes
minute set about 1/6 from anterior margin of head; propodeum with
short rounded angle, declivity not bordered; petiole 1/4 wider behind
than long, truncate dorsally in profile, narrowing so dorsum 1/4
shorter than base
TL 2.7 mm; head rectangular 1/5 longer than wide, sides slightly
arcuate, occiput straight; scape reaching occiput; head submatt, very
densely and finely puncturate, alitrunk more shiny, gaster more like
head; pubescence more obvious; clear chestnut-brown, appendages
brown-yellow |
. |
. |
 |
Kenya
& Tanzania - jeanneli |
-- |
Propodeal declivity distinctly bordered;
petiole scale a thick scale or node |
35 |
35 |
Clypeus with feeble
median carina; brown or red, with dense whitish pilosity |
36 |
-- |
Clypeus with strong median carina;
propodeum laterally bordered; body shiny brown or yellow matt but
without dense pilosity |
37 |
- |
Clypeus with feeble
median carina; brown or red, with dense whitish pilosity |
- |
36 |
TL
3.0-3.3 mm; head only slightly elongated (HL 0.8 HW 0.7) sides slightly
convex, widest at the midpoint; clypeus short, feebly angular
anteriorly; mandibles very puncturated and less curved than other
species, 7-8 denticles subtend a short tooth, followed by 4 sharp
teeth; the concave occiput is not reached by the antennal scapes,
although those attain the mid-point of the occiput concavity; no sign
of eyes, although that may be due to the density of the puncturation;
puncturation denser than the other brown species, composed of medium
sized punctures, separated by spaces of a similar size to the
punctures, specially regularly arranged on the head; dull
reddish-brown, head lighter and more shiny, but this is hidden by the
dense recumbent white pilosity, giving a matt appearance to the rest of
the body |
Guinea
- occidentalis |
-- |
TL
2.6-2.8 mm; mandibles shiny, eyes with 1-2 facets; clypeus feebly
carinated, carina rounded, most visible anteriorly; declivity of
propodeum bordered with a distinctly concave declivity; overall shiny
but head and gaster less shiny due to dense whitish recumbent pilosity,
densest on the gaster; dark brown, appendages and apex of gaster dull
yellow |
Eastern Africa - ursa |
-- |
TL
3.5 mm; eye of 1 or no facets; mandibles with distinct teeth; petiole
scale narrows from base to apex in profile |
Madagascar
(for comparison) - madecassa |
. |
Clypeus with strong
median carina; body shiny brown or yellow mat but without dense
pilosity; propodeum laterally bordered |
. |
37 |
TL
in range 2.6-3.3 mm; differs from jeanneli in structure of
clypeus and thorax; head more shiny and eyeless; clypeus with distinct
medial carina, mandibles with 3-5 teeth; body without flattened
pilosity; brown |
|
. |
 |
East
Africa & Guinea - abyssinica |
-- |
TL
2.8 mm; head elongated, sides relatively straight and parallel,
eyeless, occiput near rectilinear; clypeus longer than coeca,
with strong carina, rounded anteriorly; mandibles similar to coeca,
with 4-5 large teeth, following 8-9 denticles, a little puncturated;
head densely covered with fine puncturations, pilosity feeble; shiny,
uniform yellow, funiculus darker
(B&F synonym of occidentalis)
|
Guinea &
Cameroun - intermedia |
-- |
TL
ca 3.5 mm; petiole scale with anterior and posterior faces in profile
completely parallel; petiole apex higher than the level of the
propodeum dorsum; propodeum declivity shallowly concave |
South
Africa - new species - hlavaci |